The cup well and truely caught on in the North in 1878 as clubs from Sheffield, Darwen, Nottingham and Manchester all entered in a bid to take the cup out of the home counties. Queen's Park as usual represented Scotland and as usual they withdrew without playing a fixture. Wales was also represented for the first time as the famed Druids club lined up in round one. The home County old boys would still dominate however and the holders, Wanderers were in no mood to relinquish their hold on the cup. Panthers and High Wycombe were both hit for nine goals each by the holders in the opening two rounds with Hubert Heron netting four and Henry Wace getting a hat-trick against Panthers. A replay victory over Barnes gave the holders one of the six available quarter final spots to maintain a record of never having failed to reach that stage of the competition.
Last year's beaten finalists, Oxford University were also in fine scoring form and after a 5-2 win over Herts Rangers in round one they too booked a quarter final spot with wins over Old Forresters and Clapham.
Round ones biggest winners were Maidenhead, notching up double figures in a 10-0 win over Reading. Their own cup run came to an end at Cambridge University. The University side then met Old Harrovians, 6-0 winners over 1st Surrey Rifles in round two, for a quarter final spot which the Harrovians claimed in a replay.
Hawks had fun in what was the clubs very first cup tie when they beat Minerva 5-2 but they also went down in round two to a team curiously called Remnants. Remnants in turn failed to make the quarter final line up as they lost at Upton Park in round three.
R B Hedley had a great cup run with Royal Engineers scoring a hat-trick in a 6-0 round two win over Pilgrims and then notching another against Druids in an 8-0 romp which booked their quarter final spot. Sheffield made up the last six as the only side North of Watford still in the cup.
The Northern challenge died on quarter final day as Wanderers saw off Sheffield by 3-0. Old Harrovians defeated Upton Park 3-1 to reach their first ever semi final but in the tie of the round Oxford and Royal Engineers battled out a marathon tie which went to a second replay before the Engineers won 4-2.
With only three semi finalists there was a bye straight to the final on offer and it went to Wanderers leaving Royal Engineers as red hot favourites to book the other final place. Naturally Old Harrovians had other ideas but the Engineers were too strong on the day and came through 2-1 with the winner coming from C B Mayne.